- filtered for Croatia
Images and Species Descriptions
Text and photographs © 2011 Jørgen Lissner
The Spiders of Europe and Greenland
Family: Gnaphosidae (Ground Spiders)
Biology: The Gnaphosidae is the seventh largest spider family of the world encompassing ca. 2000 species in 8 subfamilies and 114 genera. They range in size from small to large (2-18 mm body size). The species are free-living found mailnly at ground level in open habitats but some species are house spiders, for example Scotophaeus blackwalli in the northern parts of its distributional range. They build a tubular retreat, from which they leave at night to hunt prey. The female guard her papery egg sack hidden in small holes under logs, stones, etc. Most species are active at night, however spiders of the somewhat atypical genus Micaria are diurnal hunters running rapidly about in bright sunshine.
Characters of family: The gnaphosids are rather stout ecribellate spiders with a flattened and elongate abdomen. The carapace is ovoid and rather low being smoothly convex and with a distinct fovea in most species. The head is not sharply set off from the thoracic region. Gnaphosids are fairly easily recognized by their cylindrical and parallel spinners, the anterior pair being slightly longer, and more heavily sclerotized than the posterior pair. The anterior spinners are separated from each other by approximately one spinner diameter with some exceptions, e.g. Micaria in which genus anterior spinners are closer. They have 8 eyes in 2 rows. The posterior medial eyes are often not round, but oval, triangular or reduced to slits. All eyes are with a silvery sheen except for the anterior medials, which are dark. The sternum is ovoid, pointed posteriorly. The chelicerae are robust, and the fang furrows are provided with teeth. The retromargin may have a sclerotized lamina (flat, keel-like plate) in place of teeth. This lamina is serrated in some genera. The curvature of the posterior row of eyes and the position and shape of cheliceral lamina and teeth are important characters when keying gnaphosids to genus level under the stereomicroscope. The endites usually have an oblique or transverse depression. They are provided with a serrula (row or cluster of tiny teeth on the front margin). Gnaphosids are also characterized by having 2 tarsal claws, claw tufts, and scopulae. Legs are stout and in some species there are small brushes of more and less stiff hairs present distally on metatarsus IV. Female palp is furnished with small spines and a finely toothed claw. The abdomen is often provided with dense coverage of short sleek hairs giving the abdomen a mousy-like appearance. Sometimes erect, curved setae are present, particularly at the anterior edge (see for example images of Gnaphosa lucifuga). Many species are uniformly coloured in greyish-brown or blackish colours. However, abdomens of some species have striking white patterns of spots or lines while abdomens of others are iridescent. Most males have a scutum at the anterior end. The spiracle is situated close to the spinners. Gnaphosids are entelegyne spiders often having rather large epigynes with sclerotized structures. They are somewhat variable and closely related species may be difficult to identify. Male palps are usually provided with a large tibial apophysis and the shape of this is important when identifying the species.
Genus: Aphantaulax Simon, 1878
Aphantaulax cincta (L. Koch, 1866)
Range: Croatia, Czech Republic, France (Corsica), France (Mainland), Greece (Mainland), Hungary, Italy (Mainland), Italy (Sardinia), Italy (Sicily), Malta, Portugal (Mainland), Romania, Slovakia, Spain (Mainland), Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe, North Africa, Israel (Platnick 10.0).
Aphantaulax trifasciata (O. P.-Cambridge, 1872)
Range: Andorra, Bulgaria, Croatia, France (Corsica), France (Mainland), Greece (Dodecanese Islands), Greece (Mainland), Hungary, Italy (Mainland), Italy (Sardinia), Italy (Sicily), Portugal (Mainland), Romania, Russia (Central European), Russia (Southern European), Slovakia, Spain (Balearic Islands), Spain (Mainland), Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Female.
Subadult female.
Female.
Female.
Genus: Callilepis Westring, 1874
Callilepis nocturna (Linnaeus, 1758)
Range: Andorra, Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Italy (Mainland), Italy (Sicily), Latvia, Macedonia, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Slovakia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Female.
Female.
Female.
Female.
Female.
Callilepis schuszteri (Herman, 1879)
Range: Austria, Bulgaria, Croatia, Czech Republic, France (Mainland), Germany, Greece (Mainland), Hungary, Italy (Mainland), Liechtenstein, Romania, Slovakia, Slovenia, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Genus: Drassodes Westring, 1851
Characters of genus: The abdomen has dense short hairs making the spiders appear mousy. These spiders are mostly reddish-brown to greyish-brown, and resemble spiders from the Clubionidae. However, the spinners are tubular as characteristic of the Gnaphosidae. The members of the genus are distinguished from other gnaphosids by the deeply notched trochanters. The posterior medial eyes are oval. Males have no scutum.
Drassodes cupreus (Blackwall, 1834)
Description: Very similar to D. lapidosus. The cephalothorax has a dark borderline. Size: 9-18 mm. Maturity: Females throughout year, males in spring and summer. Habitat: Dry places, such as heather, gravel beds, dunes and dry grassland. Spiders can be found in their retreats by turning stones or searching leaf litter, moss, grass tussucks etc. Range: Andorra, Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Hungary, Ireland, Italy (Mainland), Liechtenstein, Macedonia, Malta, Netherlands, Norway (Mainland), Romania, Russia (Kaliningrad Region), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Female.
Female.
Female.
Male.
Male.
Male.
Male abdomen.
Male.
Drassodes lapidosus (Walckenaer, 1802)
Size: 9-18 mm. Maturity: Females throughout year, males in spring and summer. Range: Andorra, Austria, Belgium, Bosnia/Herzegowina, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Channel Islands), Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Italy (Sardinia), Italy (Sicily), Latvia, Liechtenstein, Macedonia, Malta, Moldova, Norway (Mainland), Poland, Portugal (Azores), Portugal (Madeira), Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (Southern European), Slovakia, Slovenia, Spain (Canary Islands), Spain (Mainland), Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Male.
Male.
Male.
Female.
Female.
Female abdomen.
Female.
Female with egg cocoon.
Female.
Female.
Female.
Female.
Female.
Female.
Female.
Female.
Female.
Male.
Male.
Male.
Female.
Drassodes lutescens (C. L. Koch, 1839)
Range: Bulgaria, Croatia, France (Corsica), France (Mainland), Greece (North Aegean Islands), Italy (Mainland), Italy (Sardinia), Italy (Sicily), Portugal (Madeira), Portugal (Selvagens Islands), Russia (Central European), Russia (Eastern European), Russia (Southern European), Slovenia?, Spain (Canary Islands), Spain (Mainland), Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Mediterranean to Pakistan (Platnick 10.0).
Female.
Female abdomen.
Female abdomen with parasitic mites.
Female.
Male.
Male.
Female.
Female abdomen.
Female.
Drassodes montenegrinus (Kulczynski, 1897)
Range: Croatia, Yugoslavia (van Helsdingen 2009.1). Global range: Croatia, Serbia (Platnick 10.0).
Drassodes pubescens (Thorell, 1856)
Description: Similar in general appearance to the other two species of the genus, however slightly paler. The relatively small size of the adults ease species identification. Size: Female 6-8 mm; male 5-7 mm. Maturity: Spring and early summer. Habitat: Dry or somewhat moist habitats such as heather, forest edges and open forests. Range: Andorra, Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Greece (Crete), Greece (Dodecanese Islands), Greece (Mainland), Hungary, Italy (Mainland), Italy (Sicily), Latvia, Liechtenstein, Lithuania, Macedonia, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Female.
Female.
Female.
Female.
Female.
Female.
Drassodes rhodanicus Simon, 1914
Range: Croatia, France (Mainland) (van Helsdingen 2009.1). Global range: France (Platnick 10.0).
Drassodes striatus (L. Koch, 1866)
Range: Croatia, Hungary, Romania, Ukraine?, Yugoslavia (van Helsdingen 2009.1). Global range: Balkans (Platnick 10.0).
Genus: Drassyllus Chamberlin, 1922
Characters of genus: Posterior medial eyes larger than laterals and in most species oval and oblique. Posterior row of eyes slightly procurved and the medials are closer to each other than to the laterals. Distal end of metatarsi III og IV is furnished with a ventral comb. Mostly dark species resembling species of the Zelotes genus.
Drassyllus lutetianus (L. Koch, 1866)
Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Lithuania, Macedonia, Moldova, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Kazakhstan (Platnick 10.0).
Male.
Female.
Drassyllus praeficus (L. Koch, 1866)
Description: Very dark brown or black species with contrasting yellow-orange branchial operculae. Size: Female 5-6 mm; male 4.5-5 mm. Maturity: Spring and Summer. Habitat: Appears to be mainly coastal in Denmark. Seems to prefer dry areas with low vegetation and stones. Range: Andorra, Austria, Belgium, Bosnia/Herzegowina, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Greece (Crete), Greece (Mainland), Hungary, Italy (Mainland), Italy (Sicily), Latvia, Liechtenstein, Lithuania, Macedonia, Moldova, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Central Asia (Platnick 10.0).
Female.
Male.
Male.
Male.
Drassyllus pusillus (C. L. Koch, 1833)
Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Channel Islands), Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Malta, Moldova, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Male.
Male.
Female.
Female.
Female, note pale metatarsi og tarsi.
Drassyllus villicus (Thorell, 1875)
Range: Austria, Belgium, Bulgaria, Croatia, Czech Republic, France (Mainland), Germany, Hungary, Italy (Mainland), Macedonia, Portugal (Mainland), Romania, Slovakia, Slovenia, Spain (Mainland), Switzerland, Yugoslavia (van Helsdingen 2009.1). Global range: Europe (Platnick 10.0).
Genus: Gnaphosa Latreille, 1804
Characters of genus: Most species with dark brown cephalothorax. The abdomes is either dark brown, greyish-black or black, thickly covered with grey hairs. The genus is characterized by having the posterior row of eyes recurved.
Gnaphosa alacris Simon, 1878
Range: Andorra, Croatia, France (Corsica), France (Mainland), Italy (Mainland), Italy (Sicily), Spain (Mainland) (van Helsdingen 2009.1). Global range: France, Italy, Croatia, Morocco (Platnick 10.0).
Subadult male.
Subadult male.
Subadult male.
Male.
Male.
Male.
Female.
Female.
Male.
Male.
Female.
Female.
Gnaphosa bicolor (Hahn, 1833)
Range: Austria, Belarus, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Greece (Mainland), Hungary, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Slovakia, Slovenia, Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Georgia (Platnick 10.0).
Male.
Exuvium.
Juvenile.
Female.
Female.
Female.
Female.
Gnaphosa dolosa Herman, 1879
Range: Bulgaria, Croatia, Cyprus?, France (Mainland), Greece (Crete), Italy (Mainland), Macedonia, Romania, Russia (Eastern European), Russia (Southern European), Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Gnaphosa lucifuga (Walckenaer, 1802)
Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, France (Mainland), Germany, Greece (Mainland), Hungary, Italy (Mainland), Italy (Sardinia), Italy (Sicily), Latvia, Macedonia, Moldova, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Male.
Male.
Male.
Male.
Male.
Male.
Male.
Male.
Gnaphosa lugubris (C. L. Koch, 1839)
Range: Andorra, Austria, Belgium, Bulgaria, Croatia, Czech Republic, Estonia, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Italy (Mainland), Latvia, Liechtenstein, Macedonia, Malta, Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (Southern European), Slovakia, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Central Asia (Platnick 10.0).
Female.
Female abdomen.
Female (one eye is missing).
Female.
Female.
Gnaphosa modestior Kulczynski, 1897
Range: Bulgaria, Croatia, Hungary, Italy (Mainland), Macedonia, Romania, Slovakia, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Eastern Europe to Azerbaijan (Platnick 10.0).
Gnaphosa zeugitana Pavesi, 1880
Range: Croatia, Spain (Balearic Islands) (van Helsdingen 2009.1). Global range: North Africa (Platnick 10.0).
Genus: Haplodrassus Chamberlin, 1922
Characters of genus: Carapace elongate and fairly low. The brownish cephalothorax have complex black lines. Head much wider than both row of eyes. The posterior medial eyes are larger than the laterals as well as oval and oblique. Posterior row of eyes has the medials closer to each other than to the laterals. Anterior row of eyes straight or slightly recurved while posterior row is procurved. The distance between the outer edges of anterior medial eyes is about the same as the distance between outer edges of posterior medials. Clypeus low about equal to the distance of one anterior medial eye. Fovea short as in the Clubionidae. Chelicerae strong with lateral condyles. Chelicerae with teeht both on promargin and retromargin. Labium longer than wide and angulate where sides meets posterior border.Sternum does not extend between coxae IV. Trochanters smooth. The distal end of metatarsi III and IV is not furnished with a preening comb. Males do not have a scutum. Some species have vague patterns of chevrons on the abdomen, but these patterns are not reliable in separating the species since the variation within species is considerable. Male palp with a tibial apophysis, the shape of which is important for identification. Sometimes the shape is discernible with a lens allowing for identification in the field. The epigyne is often large and females of some species are also identifiable in the field using a lens.
Haplodrassus dalmatensis (L. Koch, 1866)
Range: Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Greece (Crete), Greece (Mainland), Hungary, Italy (Mainland), Italy (Sardinia), Italy (Sicily), Lithuania, Macedonia, Netherlands, Poland, Portugal (Madeira), Portugal (Mainland), Portugal (Selvagens Islands), Romania, Russia (Eastern European), Russia (Southern European), Slovakia, Spain (Balearic Islands), Spain (Canary Islands), Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Male.
Male.
Male.
Female.
Female abdominal markings.
Female.
Female.
Female.
Female.
Female.
Haplodrassus minor (O. P.-Cambridge, 1879)
Description: Resemble H. signifer but is much smaller. The size of adults is helpful in identifying the species. Size: Female 4 mm; male 3.5 mm. Maturity: Summer. Habitat: Coastal shingle and sandy beaches with upwash. Range: Austria, Bulgaria, Croatia, Czech Republic, Denmark, Germany, Great Britain (Mainland), Greece (Crete), Greece (Mainland), Hungary, Italy (Mainland), Macedonia, Moldova, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Southern European), Slovakia, Spain (Canary Islands), Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe, Russia (Platnick 10.0).
Male.
Male.
Male.
Male.
Male.
Male.
Male.
Male.
Male.
Female.
Female.
Female.
Haplodrassus signifer (C. L. Koch, 1839) - Stealthy ground spider
Description: Carapace greyish brown to dark brown, head gradually darker towards anterior edge in lighter specimens. Chelicerae dark brown. Sternum dark brown. Legs brown. The abdomen is somewhat flattened, brown to blackish. Three pairs of longitudinal short stripes (sigilla?) are sometimes visible on dorsum, the posterior pair oblique. Light, obscure chevrons sometimes present in lighter, mostly female specimens. Males generally darker than females, sometimes nearly black. Size: Female 7-9 mm; male 5-8 mm. Maturity: Spring and early summer. Habitat: Open, mostly dry habitats such as heathland, dunes and sandy grassland. Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Faroe Islands, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Channel Islands), Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Crete), Greece (Cyclades), Greece (Dodecanese Islands), Greece (Mainland), Hungary, Iceland, Ireland, Italy (Mainland), Italy (Sardinia), Italy (Sicily), Latvia, Liechtenstein, Lithuania, Macedonia, Malta, Moldova, Netherlands, Norway (Mainland), Poland, Portugal (Azores), Portugal (Madeira), Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Canary Islands), Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Also known from Greenland (Masurik in prep.). Global range: Holarctic (Platnick 10.0).
Subadult female.
Male.
Female before oviposition.
Female.
Male.
Male.
Male.
Juvenile.
Juvenile.
Male.
Male.
Male.
Male.
Male.
Female.
Female.
Female.
Female.
Female.
Female.
Female.
Female.
Genus: Kishidaia Yaginuma, 1960
Kishidaia conspicua (L. Koch, 1866)
Range: Austria, Belgium, Bulgaria, Croatia, Czech Republic, France (Mainland), Germany, Hungary, Italy (Mainland), Malta, Moldova, Netherlands, Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Central Asia (Platnick 10.0).
Genus: Leptodrassus Simon, 1878
Leptodrassus croaticus Dalmas, 1919
Range: Croatia (van Helsdingen 2009.1). Global range: Croatia (Platnick 10.0).
Leptodrassus femineus (Simon, 1873)
Range: Bulgaria, Croatia, France (Corsica), France (Mainland), Greece (Crete), Italy (Sicily), Portugal (Mainland) (van Helsdingen 2009.1). Global range: Portugal to Crete (Platnick 10.0).
Female.
Female.
Female.
Genus: Micaria Westring, 1851
Characters of genus: This genus has previously been assigned to the Clubionidae by some workers. A depression across the endites and the often oval posterior medial eyes has resulted in its present status as a gnaphosid genus. Tne cephalothorax and abdomen is covered with flattened dark scales, usually iridecent and sometimes with contrasting white spots or stripes. Slim, antlike spiders running rapidly over the ground in dry and warm places.
Micaria albovittata (Lucas, 1846)
Range: Andorra, Austria, Belgium, Bulgaria, Croatia, Czech Republic, France (Corsica), France (Mainland), Great Britain (Mainland), Greece (Crete), Greece (Dodecanese Islands), Greece (Mainland), Hungary, Italy (Mainland), Latvia, Macedonia, Portugal (Madeira), Portugal (Mainland), Romania, Russia (Northern European)?, Russia (NW. European)?, Slovakia, Slovenia, Spain (Mainland), Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Female.
Female.
Micaria coarctata (Lucas, 1846)
Range: Bulgaria, Croatia, France (Corsica), France (Mainland), Great Britain (Channel Islands), Greece (Crete), Greece (Cyclades), Greece (Mainland), Italy (Mainland), Malta, Portugal (Mainland), Romania, Spain (Mainland), Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Mediterranean to Central Asia (Platnick 10.0).
Female.
Female.
Female.
Female spinners.
Female with prey.
Female.
Female.
Female.
Female.
Female.
Female.
Female.
Female.
Micaria dives (Lucas, 1846) - Jewel ant-spider
Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, France (Corsica), France (Mainland), Germany, Greece (Crete), Greece (Mainland), Hungary, Italy (Mainland), Macedonia, Netherlands, Portugal (Mainland), Romania, Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Ukraine (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Juvenile.
Juvenile.
Juvenile (subadult?). Missing right foreleg regenerated almost completely in just one moult.
Subadult female.
Subadult female.
Subadult female.
Subadult female.
Female.
Female.
Female.
Male.
Male.
Male.
Male.
Male.
Male.
Micaria formicaria (Sundevall, 1831)
Range: Andorra, Austria, Belgium, Bulgaria, Croatia, Czech Republic, Estonia?, Finland, France (Mainland), Germany, Hungary, Italy (Mainland), Liechtenstein, Macedonia, Malta, Moldova, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Slovakia, Slovenia, Spain (Balearic Islands), Spain (Mainland), Sweden, Switzerland, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Subadult female.
Subadult female.
Subadult female.
Female.
Female.
Female.
Female.
Micaria fulgens (Walckenaer, 1802) - Purple ant-spider
Size: Female 4.5-6 mm; male 4.5-5.5 mm. Range: Andorra, Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Greece (Mainland), Hungary, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (NW. European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Central Asia, North Africa (Platnick 10.0).
Female.
Female.
Female.
Female.
Micaria pulicaria (Sundevall, 1831) - Glossy ant-spider
Description: Head black with radiating white lines. Iridescense less pronounced compared to the highly iredescent abdomen. There are two transverse, white lines on the abdomen, followed by three dots. Femora I and II are black, while femora III and IV are dark brown. Size: Female 2.7-4.5 mm; male 3-3.5 mm. Maturity: Spring and summer. Habitat: Dry and sunny places in a variety of situations, including gardens. Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Channel Islands), Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Crete), Greece (Mainland), Hungary, Iceland, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Spain (Mainland), Sweden, Switzerland, Yugoslavia (van Helsdingen 2009.1). Global range: Holarctic (Platnick 10.0).
Female.
Male.
Female.
Female.
Female.
Female.
Micaria sociabilis Kulczynski, 1897
Range: Bulgaria, Croatia, Czech Republic, France (Corsica), France (Mainland), Hungary, Italy (Mainland), Macedonia, Portugal (Mainland), Romania, Slovakia, Spain (Mainland), Yugoslavia (van Helsdingen 2009.1). Global range: Central, Southern Europe (Platnick 10.0).
Genus: Nomisia Dalmas, 1921
Nomisia aussereri (L. Koch, 1872)
Range: Bulgaria, Croatia, France (Corsica), France (Mainland), Greece (Cyclades), Greece (Dodecanese Islands), Greece (Mainland), Greece (North Aegean Islands), Italy (Mainland), Italy (Sardinia), Italy (Sicily), Macedonia, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Southern European), Spain (Mainland), Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Nomisia exornata (C. L. Koch, 1839)
Range: Bulgaria, Croatia, France (Corsica), France (Mainland), Greece (Crete), Greece (Cyclades), Greece (Dodecanese Islands), Greece (Mainland), Greece (North Aegean Islands), Hungary, Italy (Mainland), Italy (Sardinia), Italy (Sicily), Macedonia, Malta, Portugal (Mainland), Romania, Slovakia, Spain (Balearic Islands), Spain (Mainland), Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Central Asia (Platnick 10.0).
Male.
Male.
Male.
Female.
Female.
Female.
Female.
Female.
Female.
Female.
Female, abdominal markings.
Female.
Female.
Nomisia molendinaria (L. Koch, 1866)
Range: Croatia, Italy (Mainland) (van Helsdingen 2009.1). Global range: Croatia, Georgia (Platnick 10.0).
Genus: Phaeocedus Simon, 1893
Phaeocedus braccatus (L. Koch, 1866)
Range: Andorra, Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Hungary, Italy (Mainland), Italy (Sicily), Latvia, Macedonia, Netherlands, Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (NW. European)?, Russia (Southern European), Slovakia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Female.
Female.
Female.
Female.
Female.
Genus: Scotophaeus Simon, 1893
Characters of genus: This genus has three species in Northern Europe of which two occur in Denmark. The abdomen is without a pattern and appear greasy silverish-grey due to the short, dense grey hairs. The males have a conspicious brown abdominal scutum, but is small and therefore not clearly visible for the commonest species of the genus, S. blackwalli. The posterior medial eyes are circular and slightly closer to another than to the laterals. Height of clypeus about the same as the diameter of an anterior medial eye. In Northern Europe the species are found exclusively within houses where they wander about at nighttime.
Scotophaeus blackwalli (Thorell, 1871)
Description: This species is also known under the name Herpyllus blackwalli. Males slightly smaller than females, otherwise the sexes are similar. The cephalothorax and legs are orange brown covered with grey hairs. Size: Female 9-12 mm; male 7-10 mm. Maturity: Females throughout year, males in summer and autumn. Habitat: In Denmark only inside heated houses. In warmer parts of Europe also outdoors under bark and crevices in walls. Range: Austria, Belgium, Bulgaria, Croatia, Denmark, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Greece (Dodecanese Islands), Hungary, Ireland, Italy (Mainland), Italy (Sicily), Malta, Netherlands, Norway (Mainland), Poland, Portugal (Azores), Portugal (Madeira), Portugal (Mainland), Portugal (Selvagens Islands), Romania, Russia (Northern European), Slovakia, Spain (Canary Islands), Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Cosmopolitan (Platnick 10.0).
Female.
Female.
Subadult female.
Typical cylindrical Gnaphosid spinnerets.
Female.
Female abdomen.
Female.
Female.
Male.
Male.
Female.
Male.
Male.
Scotophaeus blackwalli isabellinus (Simon, 1873)
Range: Croatia, France (Corsica), Portugal (Mainland) (van Helsdingen 2009.1). Global range: Corsica, Italy, Croatia (Platnick 10.0).
Scotophaeus quadripunctatus (Linnaeus, 1758)
Range: Austria, Belarus, Bulgaria, Croatia, Czech Republic, Estonia, France (Mainland), Germany, Hungary, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Moldova, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (NW. European), Russia (Southern European), Slovakia, Sweden, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Europe, Russia (Platnick 10.0).
Scotophaeus scutulatus (L. Koch, 1866)
Range: Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, France (Mainland), Germany, Great Britain (Mainland), Greece (Crete), Greece (Cyclades), Greece (Mainland), Hungary, Italy (Mainland), Italy (Sardinia), Liechtenstein, Lithuania?, Macedonia, Malta, Moldova, Netherlands, Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Central Asia, Algeria (Platnick 10.0).
Female.
Female.
Female.
Female abdomen.
Female.
Female.
Female.
Female.
Female.
Habitat, stem with bark peeled off to reveal eggsack .
Female.
Female.
Female, leg with scopula.
Female.
Female, abdominal markings.
Male.
Male.
Male, note that scutum is not visible on living specimens.
Female.
Female.
Genus: Setaphis Simon, 1893
Setaphis carmeli (O. P.-Cambridge, 1872)
Range: Bulgaria, Croatia, France (Corsica), France (Mainland), Greece (Crete), Greece (Dodecanese Islands), Greece (Mainland), Italy (Mainland), Italy (Sardinia), Italy (Sicily), Macedonia, Portugal (Mainland), Spain (Balearic Islands), Spain (Mainland) (van Helsdingen 2009.1). Global range: Mediterranean (Platnick 10.0).
Genus: Trachyzelotes Lohmander, 1944
Trachyzelotes adriaticus (Caporiacco, 1951)
Range: Croatia, Cyprus, Greece (Crete), Greece (Mainland), Italy (Mainland), Italy (Sicily), Russia (Eastern European), Russia (Southern European) (van Helsdingen 2009.1). Global range: Italy to China (Platnick 10.0).
Female.
Trachyzelotes barbatus (L. Koch, 1866)
Range: Bulgaria, Croatia, France (Corsica), France (Mainland), Greece (Crete), Greece (Mainland), Italy (Mainland), Italy (Sicily), Malta, Portugal (Mainland), Romania, Russia (Central European), Russia (Southern European), Slovenia?, Spain (Balearic Islands), Spain (Mainland) (van Helsdingen 2009.1). Global range: Mediterranean to Central Asia, USA (Platnick 10.0).
Male.
Male.
Male.
Male.
Female.
Female.
Female.
Female.
Female.
Trachyzelotes fuscipes (L. Koch, 1866)
Range: Croatia, France (Corsica), France (Mainland), Greece (Dodecanese Islands), Italy (Mainland), Italy (Sicily), Portugal (Mainland), Spain (Mainland) (van Helsdingen 2009.1). Global range: Mediterranean, China (Platnick 10.0).
Female.
Trachyzelotes holosericeus (Simon, 1878)
Range: Croatia, France (Mainland), Greece (Mainland), Portugal (Madeira), Portugal (Mainland), Spain (Mainland) (van Helsdingen 2009.1). Global range: Western Mediterranean (Platnick 10.0).
Male.
Male.
Male.
Trachyzelotes lyonneti (Audouin, 1826)
Range: Croatia, France (Mainland), Greece (Crete), Greece (North Aegean Islands), Italy (Mainland), Italy (Sicily), Portugal (Azores), Portugal (Madeira), Russia (Southern European), Spain (Canary Islands), Ukraine (van Helsdingen 2009.1). Global range: Mediterranean to Central Asia, USA, Brazil, Peru (Platnick 10.0).
Female.
Palp.
Male.
Female.
Female.
Female.
Trachyzelotes pedestris (C. L. Koch, 1837)
Range: Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, France (Mainland), Germany, Great Britain (Mainland), Greece (Dodecanese Islands), Greece (Mainland), Hungary, Italy (Mainland), Italy (Sicily), Latvia, Liechtenstein, Macedonia, Netherlands, Poland, Portugal (Mainland), Romania, Russia (Kaliningrad Region), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Azerbaijan (Platnick 10.0).
Male.
Male.
Female.
Female.
Female.
Female.
Female.
Female.
Female.
Genus: Urozelotes Mello-Leitão, 1938
Urozelotes rusticus (L. Koch, 1872)
Range: Austria, Bulgaria, Croatia, Czech Republic, Denmark (introduced), France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Italy (Mainland), Italy (Sicily), Macedonia, Portugal (Madeira), Portugal (Mainland), Portugal (Selvagens Islands), Romania, Russia (Central European), Russia (NW. European), Russia (Southern European), Spain (Canary Islands) (introduced), Spain (Mainland), Switzerland (van Helsdingen 2009.1). Global range: Cosmopolitan (Platnick 10.0).
Genus: Zelotes Gistel, 1848
Characters of genus: Brownish-black to black spiders without markings. Resemble dark species of Drassyllus. Posterior medial eyes are of about the the same size as the laterals. Posterior eyerow is straight, rarely procurved and eyes equidistant. The eye rows are short due to a narrow head region, the width of an eyerow less than one third of the width of the carapace at its widest point. The carapace is usually shiny and shinier than the abdomen. Some species of the genus have been moved to other genera probably because it used to be species rich with more than fifty species known from France. Several species are almost identical and needs microscopic examination of the genitialia to be separated with certainity. Some species may in some cases be identified by the markings on the legs.
Zelotes apricorum (L. Koch, 1876)
Range: Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia?, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Italy (Sicily), Latvia?, Liechtenstein, Macedonia, Moldova, Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Kazakhstan (Platnick 10.0).
Female, wet with dew.
Female, wet with dew.
Zelotes atrocaeruleus (Simon, 1878)
Range: Austria, Bulgaria, Croatia, Czech Republic, France (Mainland), Germany, Hungary, Italy (Mainland), Italy (Sicily), Macedonia, Romania, Russia (Eastern European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Switzerland (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Zelotes caucasius (L. Koch, 1866)
Range: Austria, Bulgaria, Croatia, Czech Republic, France (Mainland), Greece (Crete), Greece (Dodecanese Islands), Greece (Mainland), Hungary, Italy (Mainland), Italy (Sicily), Macedonia, Portugal (Mainland), Romania, Russia (Eastern European), Russia (Southern European), Slovakia, Spain (Mainland), Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Central Asia (Platnick 10.0).
Female.
Female.
Male.
Male.
Male.
Male.
Zelotes clivicola (L. Koch, 1870)
Range: Austria, Belarus, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Greece (Crete), Greece (Mainland), Hungary?, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Male.
Male.
Zelotes electus (C. L. Koch, 1839)
Description: Zelotes electus has stout legs compared to other Zelotes species and therefore appear robust. The species is also characterised by compactly grouped eyes, where the anterior medials are contiguous with the anterior laterals. The pigmentation of the legs aids in distinguishing the species in this genus as well as for a few closely related genera. Zelotes electus has femora brown but darkened apically, dark patella, dark tibia, brown or orange metatarsi and tarsi. Size: Female 4-5.5 mm; male 3.5-4.5 mm. Maturity: Spring and summer. Habitat: Locally common in coastal dunes with lichens and low vegetation sometimes also further inland. The spider is frquently seen running about in sunshine. Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Dodecanese Islands)?, Greece (Mainland), Hungary, Ireland, Italy (Mainland), Italy (Sicily), Lithuania, Macedonia, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Central Asia (Platnick 10.0).
Female.
Female.
Female.
Female.
Female.
Female.
Zelotes erebeus (Thorell, 1871)
Range: Austria, Belgium, Bulgaria, Croatia, Czech Republic, France (Mainland), Germany, Greece (Mainland), Hungary, Italy (Mainland), Macedonia, Portugal (Mainland), Romania, Russia (Eastern European), Slovakia, Slovenia, Switzerland, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Georgia (Platnick 10.0).
Male.
Male.
Male.
Male.
Female (Photo © Dimitri Geystor, FR).
Female (Photo © Dimitri Geystor, FR).
Epigyne (Photo © Dimitri Geystor, FR).
Zelotes femellus (L. Koch, 1866)
Range: Croatia, France (Corsica), Greece (Mainland), Italy (Mainland), Italy (Sardinia), Italy (Sicily), Romania, Spain (Mainland), Ukraine (van Helsdingen 2009.1). Global range: Southern Europe (Platnick 10.0).
Zelotes gracilis (Canestrini, 1868)
Range: Austria, Bulgaria, Croatia, Czech Republic, Hungary, Italy (Mainland), Macedonia, Moldova, Romania, Russia (Southern European), Slovakia, Slovenia, Ukraine (van Helsdingen 2009.1). Global range: Europe, Russia (Platnick 10.0).
Zelotes hermani (Chyzer, 1897)
Range: Austria, Bulgaria, Croatia, Hungary, Italy (Mainland), Macedonia, Romania, Slovakia, Slovenia, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Central Europe to Russia (Platnick 10.0).
Zelotes latreillei (Simon, 1878)
Description: Large, shiny black species. Except for orange operculae, the only non black parts are the tarsi which are brownish-grey. Size: Female 7-8 mm; male 4.5-7.5 mm. Maturity: Spring and summer. Habitat: In a variety of dry situations, such as dunes, sandy foreland and open forest on sandy soil. Range: Andorra, Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Moldova, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Female.
Female.
Zelotes oblongus (C. L. Koch, 1833)
Range: Austria, Bulgaria, Croatia, France (Mainland), Germany, Greece (Crete), Greece (Mainland), Italy (Mainland), Macedonia, Romania, Switzerland, Yugoslavia (van Helsdingen 2009.1). Global range: Europe (Platnick 10.0).
Zelotes similis (Kulczynski, 1887)
Range: Austria, Bulgaria, Croatia, Germany, Greece (Mainland), Hungary, Italy (Mainland), Liechtenstein, Romania, Switzerland, Yugoslavia (van Helsdingen 2009.1). Global range: Europe (Platnick 10.0).
Zelotes tenuis (L. Koch, 1866)
Range: Austria, Bulgaria, Croatia, France (Corsica), France (Mainland), Greece (Crete), Greece (Dodecanese Islands), Greece (Mainland), Italy (Mainland), Italy (Sicily), Macedonia, Malta, Portugal (Azores), Portugal (Mainland), Romania, Spain (Canary Islands), Spain (Mainland), Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Mediterranean to Ukraine, USA (Platnick 10.0).
Palp and epigyne.
Male.
Male.
Female.